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Botticelli was closely associated with the Medici and his fortune paralleled theirs. After the death of Lorenzo, that ended the world in which Botticelli had found honours and fame, the painter was greatly impressed by the preaching of Savonarola. Soon he became an ardent disciple of this great prophet. When Savonarola demanded that bonfires should be made of the «profane pictures», he contributed many of his works of art to the bonfire pile. In his later life Botticelli turned to a religious style, and after 1500 gave up painting altogether.

Botticelli's most celebrated pictures, the Primavera (The Allegory of Spring) and the Birth of Venus were painted at a slight distance from each other in time, the first on panel, the second on canvas. Later the two paintings were considered companion pieces. Both have been interpreted in different ways. The Primavera with its ambiguous but clear meaning, is far from being the simple pagan mythology that it appears to be at first sight. No explanation of the Primavera is wholly successful. Probably the Primavera symbolizes Lorenzo Medici's real wedding in 1482.

A Christianized Venus, modestly dressed and resembling Botticelli's Madonnas, reigns in the midst of a dark grove of trees bearing golden fruit. At the right Zephyrus, the wind-god, pursues the nymph Chloris; flowers issue from her mouth. She is transformed into the goddess Flora, clothed in a flower-covered gown, from its folds she strews blossoms upon the lawn. At the left Mercury is dispelling tiny clouds from the golden apple, the symbol of the Medici family. Between Mercury and Venus the Three Graces dance in a ring. These lovely creatures are shown in transparent garments. This painting is a complex allegory. As in all Botticelli's mature works his figures are extremely attenuated, with long necks, torsos, arms and sloping shoulders. Their beautiful faces and graceful bodies and limbs seem almost bloodless and weightless, their white feet touch the ground so lightly that not a flower or a leaf is bent. The individual forms are perfectly modelled. Botticelli's representation of figures in motion is far beyond anything that preceded him and has never been excelled. The composition is based on an interweaving of linear patterns, drapery folds, streaming or braided hair, trunks, and leaves. Such a picture, both in content and style, represents a withdrawal from naturalism of the Early Florentine Renaissance.

The Birth of Venus may show the effects of Botticelli's residence in Rome in the early 1480s. Venus, according to the ancient myth, was born from the sea. Upon a sea represented without concern for space, and dotted with little V-shaped marks for waves, Botticelli's Venus stands lightly in a beautiful cockleshell, wafted by two embracing wind-gods, toward a highly stylized shore. This Venus, proportioned like the Three Graces, differs from the splendid Venuses of classical antiquity. She uses the curving streams of her long hair to cover her nakedness. She can't wait for the cloak that one of the Hours is about to spread around her. Botticelli's allegory is related to the Christian tradition with which he tried to reconcile the pagan legend. The composition has been compared to medieval and Renaissance representations of the Baptism of Christ. It may be argued that this is a rather artificial interpretation, but it is an interpretation that made sense to the fifteenth century.